[ 1] Larkin , S cow craf t .Somaclonal variati on a novel s ou rce of variabilit y f or cell cul tu res f or plant imp rovemen t [ J] .TAG ,1981 , 60 :197-214 .[ 2] 胡尚莲, 曾寒冰.小麦单细胞培养的研究———再生植株后代主要农艺性状变异及遗传稳定性[ J] .东北农业大学学报,1996 , 27(1):1-8 .
[ 3] Ryan .Somaclonal variation in some ag rin omi c and qualitych aract ers in w heat[ J] .TAG , 1987 , 74 :77-84 .
[ 4] 朱至清, 桑建利, 王历秀, 等.体细胞无性系变异培育高蛋白种质[ J] .植物学报, 1992 , 34(12):912-918 .
[ 5] 王培, 朱至清.冬小麦体细胞无性系种子蛋白质含量的遗传[ J] .华北农学报, 1995 , 10(3):6-9 .
[ 6] 刘锦红, 胡尚莲, 曾寒冰.体细胞无性系变异改良小麦品质的研究———第7 和第8 代系籽粒蛋白质和面筋含量分析[ J] .东北农业大学学报, 1997 , 28(4):334-339 .
[ 7] 刘景松, 宋天君, 孙岩, 等.小麦体细胞无性系91B569 的变异分析[ J] .黑龙江农业科学, 2006(5):17-19 .
[ 8] 王培, 范光年, 方仁, 等.幼穗无性系变异在育种上的应用[ J] .作物学报, 1992 , 18(5):391-396 .
[ 9] 张怀刚, 陈集贤,胡含.小麦体细胞无性系变异SDS 沉降值的变异[ J] .西北农学报, 1998 , 7(2):1-5 .
[ 10] 陆维忠, 姚金宝.体细胞无性系变异与小麦赤霉病抗性改良[ M] ∥陆维忠, 郑企成.植物细胞工程与分子育种技术研究.北京:中国农业科学技术出版社, 2003 :39-47 .
[ 11] Bar ro F , Rooke L , Bekes F .Transf ormati on of w heat wi thhigh molecu lar w eigh t sub urit genes res ult s in imp roved funcitional p ropert ies [ J ] .Natu re Biot achnology , 1997 , 15 :1295-1299 .
[ 12] H alford N G, Fried J M , Blai r H , et al.Analysis of HMW glutenin sub uit s encoded by chroms om el of b read w heat(t rit icumaes tivum L .)indi cates quanti tat ive eff ect s on grain quat ity[ J] .Tkeor Appl .Genet , 1992 , 83 :373-378 .
[ 13] Payne P I , Night ingat e M A, krat tinger A F, et al.The relationship betw een HMW glu tein s ubunit cam positi on and thebread making qualit y of Bri ti sh-grow n w heat variet ies [ J] .JSci Food Ag ri c, 1987 , 40 :51-56 .
[ 14] Al tpet er F, Vasil V , S ri vas tava V , et al.in tegration an d expressionof the hig h-m ol ecu lar-w eig h glut enin sub unit 1A x1gene int o w hat[ J] .Nat Biot echnol.1996 , 14(9):1155-1159 .
[ 15] Blechl A E , Aderson O D .Exp ressi on of a novel high-molecular-w ei ght glut enin su bunit gene in t ransgenic w heat[ J] .NutureBiot ech nol , 1996 , 14 :875-879 .
[ 16] Alvarez M L, Gomez M , Carrill o T M , et al.Analysi s of daughfun cti onali ty of f lours f rom t ransgenic w heat [ J] .MulecularBreeding , 2001 , 8(1):103-108 .
[ 17] Popineau Y , Deshayes G , Lef ehv re J , et al.Prolamin aggregation , gent en vascoel as licit y , and mixing properties of t ransgenetic w heat lines exp ressing 1Ax and 1Dx high molecularw eight g lut inin subu ri t transgenes[ J] .Jou rn al of A ri culturaland Food ch emi st ry , 2001 , 49(1):395-401 .
[ 18] Rooke L , Bekes F , fido R, et al.Orerexp ressi on of a gen tenprotein in t ran sgeni c w heat resu lt s in great y increased doughst reng th [ J] .J ou rnal o Cereal S cience, 1999 , 30 :115-120 .
[ 19] He G Y, Rook e L , S t eete S , et al.T ran sf orm at ion of pestaw h eat (T riti cum turgidum L.Var du ram)wi th high-molecular-w eigh t glu tenin subuni t genes and modifi cation of doughfun cti onali ty [ J] .Molecul ar Breeding , 1999 , 5 :377-386 .
[ 20] Vasi l Z K , Bean S , Zh ao J M , et al .Evalu at ion of bakin g properties and g lut en prot ein composit ion of f ield grow n transgenicw heat lines expressin g high molecular weight glut enin gen e1Ax 1 [ J] .J plant phyiol 2001 ,158 :521-528 .
[ 21] 陈梁鸿, 王新望, 张晓东, 等.小麦编码高分子量谷蛋白亚基基因转化[ J] .作物学报, 1999 , 25(4):437-440 .
[ 22] 张晓东, 梁荣奇, 陈绪清, 等.优质HMW-GS 谷蛋白亚基转基因小麦的获得及其遗传稳定性和品质性状分析[ J] .科学通报, 2003 , 48(5):474-479 .
[ 23] 王广金, 李忠杰, 张晓东, 等.利用花粉管通道法将编码优质HMW-GS 基因导入小麦进行品质改良的研究[ J] .黑龙江农业科学, 2002(6):1-3 .
[ 24] 梁静静, 吕德彬, 陈军营, 等.小麦5 +10 亚基基因转化研究[ J] .麦类作物学报, 2004 , 24(3):5-8 .
[ 25] 施农农, 何光源, 李克秀, 等.基因枪法获得优质亚基基因表达的转基因小麦[ J] .中国农业科学, 2005 , 38(5):874-881 .
[ 26] 陈军营, 义付善, 程西永, 等.农杆菌介导小麦HMW-GS(5 +10)基因转化研究[ J] .河南农业大学学报, 2006 , 40(5):459-463 .
[ 27] 孟超敏, 陈绪清, 梁奇荣, 等.高赖酸含量基因在转基小麦的表达及其赖氨酸含量分析[ J] .科学通报, 2004 , 49(7):1731-1735 .
[ 28] 苏明玉, 王雪表, 张艳锋, 等.转大豆DNA 获得小麦高蛋白材料及品质分析[ J] .郑州大学学报(理学版), 2002 , 34(3):46-49 .
[ 29] 马华平, 赵宗武, 蒋忠凯, 等.离子束介导转大豆DNA 小麦后代品质和农艺性状分析[ J] .河南农业科学, 2003(7):4-6 .
[ 30] Smith H A , Barian a H S , ogb on naya F C , et al.Implemen tation of markers in Aus tralian w heat breading [ J] .Aus t ralianJournal of Agricul toral Reseach , 2001 , 52 :1349-1356 .
[ 31] De Bust os A , Ru bio P , Soler C , et al.Markerassis ted select ionto im prove HMW-GS-glut enins in w heat [ J] .Euphy tica ,2001 , 119 :69-73 .
[ 32] De bus tos A , rubio P , Jouve N .Molecular charact erization ofth e inacti ve allele of gen Glu-A1 and the devel opm ent of a setof AS-PCK markers f or HM W gent enin of w h eat [ J] .Th eoretical and Applied Genet ics , 2000 , 100:1085-1094 .
[ 33] Zhang W, Gianibelli M C , Rampling L , et al charact eriz at ionand marker development f or low m oleculer w eigh glu teringenes f rom Glu-A3 alleles of breed w heat [ J] .Theoret ical andAppliea Gen et ices , 2004 , 108 :1409-1419 .
[ 34] Zh ang W, Gianibelli M C , Ma W, et al.Identi ficat ion of SNPSand devel opm ent allele-s pecif ic PC R m ark es for g-gliadin allelesin T riti cum aest ivum [ J] .Theoretical an d Applied Genetics, 2003 , 107 :130-138 .
[ 35] Pras ad M , Varshney R K , Kum ar A , et al .A microsatalli tsmarker-ass ociated wi th a QT L f or grain prot ein con tent onch romosome are IDL of bread wh eat [ J] .Theoret ical and AppliedGenet ics , 1999 , 99 :341-345 .
[ 36] Prasad M , Kumar N , Kulwal P L , et al.Q TL analy sis f orgrain protein cont en t using SSR markers and validat ion seudiesu sing NILS in b reed w heat [ J] .Theoret ical and AppliedGeneti es ,2003 , 106(4):659-667 .
[ 37] Gi rout M J , M orri s C F.A egycine to serine change in py roindolineb is associat ed wi th w h eat grain hard nes s and low levels of st arch-surf ace f riabilin [ J] .Th eoreti cal an d Appliedgeret ics , 1997 , 95 :857-864 .
[ 38] Morris G F .Puroindolines :the m olecu lar geneti c basis ofw heat grain bardness [ J] .plant molecular Biology , 2002 , 48 :633-647 .
[ 39] Morris G F, Lil lem o M , someone M C ,et al.prevalence of puroindolinesgrain hardness gen otypes among his torical ly significant mo rth American s pring and winter w heat s [ J] .C ropSci ence, 2001 , 41 :218-228 .
[ 40] Igrejas G , Leroy P, charmet G , et al.Mapping QT Ls for grainhardness and purind oline cont ernt in w heat (Trit ti cumaestvumL)[ J] .Theoretical and Appli ed Genetics , 2002 , 106(1):19-27 .
[ 41] Campbell K G , Bergman C J , Gualbert o D G , et al.Quant it ative t rait s loci associat et wi th kernel t rait s in a sof t and times:hard wh eat cross [ J] .Crop S cience , 1999 , 95:622-628 .
[ 42] Perret an t M R, Cadalen T , C harm et G , et al.QT L analysis ofbread making quali ty in w heat using a dou bled h apl vid population [ J ] .Theoret ical and Appied Genet ics , 2000 , 100 :1167-1173 .
[ 43] Parker G D , Chalmers K J , Rathjen A J , et al.Mapping lociassocid ted w ith mil ling yield in w heat (Trit icum aes tivum L .)[ J] .Mol ecu lar Breading , 1999 , 5 :561-568 .
[ 44] Ain sw orth C C , Clark J R, Bol sdon J .Expression , organizati onand s t ru cture of the genes encoding the way p rotein (granuleboundst arch syn th ase)in w heat [ J] .Plan t M olecu lar Bi ol ogy, 1993 , 22 :67-82 .
[ 45] Chao S , S harp P J , Worland , et al.RFLP-b ased genet ic m apsof w heat lomeol ogou s g rou p T chromosomes [ J] .Th eoreti caland Ap plied Geneti cs , 1989 , 78 :495-504.
[ 46] Nakam ura T , Vrint en P , Si t o M , et al.Rapid Clas si ficat ion ofparti al w axg w heats u sing PCK-based markers [ J] .Genome ,2002 , 45(6):1150-1156 .
[ 47] 梁荣奇, 张义荣, 唐晓晖, 等.利用Wx 基因分子标记辅助选择培养面条专用优质小麦[ J] .农业生物技术学报, 2001 , 9(13):269-273 .
[ 48] 梁荣奇, 张义荣, 姚大年, 等.小麦淀粉品质改良的综合标记辅助选择体系的建立[ J] .中国农业大学学报, 2002 , 35(33):245-249 .